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Timothy Linley

Timothy Linley

Pacific Northwest National Laboratory
PO Box 999
Richland, WA 99352


Tim Linley came to the PNNL in 2010 and joined the Ecology group the same year. He has conducted research on fish geochemistry, osmoregulation, reproduction and behavior, and acoustic telemetry. He has also managed facilities involved in various aspects of fish production, including rearing, adult harvest and brood stock management, stock enhancement, and restoration projects.

Research Interests

  • Fish geochemistry
  • Fish physiology and culture

Education and Credentials

  • Ph.D., Fisheries, University of Washington, 1993
  • M.S., Fisheries, University of Washington, 1988
  • B.S., Wildlife Ecology, University of Wisconsin, 1979

PNNL Publications


  • Arola D.D., S.C. Murcia Sanchez, M. Stossel, R. Pahuja, T.J. Linley, A. Devaraj, and M. Ramulu, et al. 2018. "The limiting layer of fish scales: structure and properties." Acta Biomaterialia 67. PNNL-SA-136535. doi:10.1016/j.actbio.2017.12.011


  • Murcia Sanchez S.C., E. Lavoie, T.J. Linley, A. Devaraj, E.A. Ossa, and D.D. Arola. 2017. "The natural armors of fish: A comparison of the lamination pattern and structure of scales." JOURNAL OF THE MECHANICAL BEHAVIOR OF BIOMEDICAL MATERIALS, 73. PNNL-SA-129872. doi:10.1016/j.jmbbm.2016.09.025


  • Geist D.R., A.H. Colotelo, T.J. Linley, K.A. Wagner, and A.L. Miracle. 2016. "Effects of a Novel Fish Transport System on the Health of Adult Fall Chinook Salmon." Journal of Fish and Wildlife Management 7, no. 2:347-358. PNNL-SA-113224. doi:10.3996/102015-JFWM-108
  • Linley T.J., E.J. Krogstad, M.K. Nims, and R.B. Langshaw. 2016. "Geochemical signatures in fin rays provide a nonlethal method to distinguish the natal rearing streams of endangered juvenile Chinook Salmon Oncorhynchus tshawytscha in the Wenatchee River, Washington." Fisheries Research 181. PNNL-SA-117248. doi:10.1016/j.fishres.2016.04.004
  • Linley T.J., E.J. Krogstad, R.P. Mueller, G.A. Gill, and B.K. Lasorsa. 2016. "Mercury concentrations in Pacific lamprey (Entosphenus tridentatus) and sediments in the Columbia River Basin." Environmental Toxicology and Chemistry 35, no. 10:2571-2576. PNNL-SA-116607. doi:10.1002/etc.3423


  • Bellgraph B.J., K.M. Carter, M.A. Chamness, T.K. Abel, T.J. Linley, and V.I. Cullinan. 2014. "Predation Susceptibility of Juvenile Fall Chinook Salmon Exposed to Sudden Temperature Changes and Slightly Supersaturated Dissolved Gas." In Snake River Fall Chinook Salmon Life History Investigations, 1/1/2012 - 12/31/2012 Annual Report, edited by KF Tiffan, et al. 61-90. Pasco, Washington:Bonneville Power Administration. PNNL-SA-96230.


  • Brown R.S., K.V. Cook, B.D. Pflugrath, L.L. Rozeboom, R.C. Johnson, J. McLellan, and T.J. Linley, et al. 2013. "Vulnerability of larval and juvenile white sturgeon to barotrauma: can they handle the pressure?." Conservation Physiology 1, no. 1:Article No. cot019. PNNL-SA-96109. doi:10.1093/conphys/cot019
  • Geist D.R., T.J. Linley, V.I. Cullinan, and Z. Deng. 2013. "The effects of total dissolved gas on chum salmon fry survival, growth, gas bubble disease, and seawater tolerance." North American Journal of Fisheries Management 33, no. 1:200-215. PNNL-SA-88149. doi:10.1080/02755947.2012.750634

Selected Publications


  • Jury, S.H., M. Betka, T. Linley, and H.W. Harris. 2005. The interactions of low pH, aluminum and calcium on olfaction of Atlantic salmon (Salmo salar). Integrative and Comparative Biology. 45(6): 1151.


  • Linley, T, 2001. Influence of short-term estuarine rearing on the ocean survival and size at return of coho salmon in southeastern Alaska. N. Am. J. Aqua. 63: 306-311.
  • Linley, T. 2001. A comparison of first feeding characteristics in two populations of Chinook salmon. Transactions of the American Fisheries Society 130:519-525.


  • Hovda, J., and T.J. Linley. 2000. The potential application of hypothermia for anesthesia in adult Pacific salmon. N. Am. J. Aqua. 62: 67-72.
  • Smoker, W.W., B.A. Bachen, G. Freitag, H.J. Geiger and T.J. Linley. 2000. Alaska ocean ranching contributions to sustainable salmon fisheries. Pages 407-420 in E.E. Knudson, C.R. Steward, D.D. MacDonald, J.E. Williams, and D.W. Reiser, editors. Sustainable Fisheries Management. CRC Press. Boca Raton.


  • Smoker, W.W. and T. Linley. 1997. Are Prince William Sound hatcheries a fools bargain? Alaska Fishery Research Bulletin. 4: 75-78.


Additional Information